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KMID : 0368419700130010033
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Journal of Plant Biology
1970 Volume.13 No. 1 p.33 ~ p.46
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Pattern and association within Pinus densiflora communities in Kyunggi Province, Korea
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Abstract
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Pinus densiflora stands are common secondary forest communitied on infertile scils in Korea. The stands are results of long severe past biotic pressure such as cutting, burning and grazing. These could be regarded as biotic climax in Korea. Because of their prevalent occurrence, relatively simple species and age composition, and their domestic economic importance, study of their distributional patterns may give some basic knowledge for better utilization of land resources in Korea.
To detect distributional patterns and interspecific associations ten pine stands, each of which was homogenious with respect to topography and physiognomy, were subjectively selected from pine stanes in Kyunggi Province near Seoul in 1969 and were made object of this study.
Four contiguous systematic samples of count for trees, shrubs and seedlings from belt transects were collected from homogeneous areas within ten natural pine stands. The belt transect was 64 m or 128 m in length, and 1 m, 2 m or 4 m in width. Basic units within the transect ranged from 64 to 256. The data from the contiguous transects were analysed in terms of multiple split-plot experiment. Departure from randomness of stem distribution, i.e., pattern, was tested in terms of variance mean ratio. For the detection of association between species, correlation coefficient was calculated for different block sizes. The values of r were tested by the usual t-test.
Pine trees within one of the stands showed significant regular distribution throughout the blocks. Within other eight stands pines were randomly distributed at basic unit with 4×4 m, 2×2 m, 2×1 m and 1×1 m. One signifcantly clumped distribution at basic unit 2×2 m, however, was observed from one of the pine stands. These randomly distributed groups were themselves significantly regularly distributed throughout the blocks for four pine stands. For the other four pine stands, in addition to the random distribution at the basie unit (the primary random group), randomly distributed groups with 32 m dimension (the secondary random groups) were also observed. Both the primary and the secondary random groups were significantly regularly distributed at the rest of blocks.
Pine seedlings were not distributed randomly thoughout the blocks. Within three of the ten stands they were contagiously distributed.
Important strub species underneath pines such as Querus serrata, Q. acutissima, Lespedza intermedia, Rhododendron Yedoense var poukhanenae, Juniperus utilis, Rhododemdron mucronulatum var. ciliatum shnwed consistently similar distributional pattern with the pine at each stand.
The shrub species pairs; Rhododemdron Yedoense var. poukhanenae/Quercus serrata, Rhododendron mucronulatum var. ciliatum/ Lsepedeza intermedia were significantly negatively associated from 1 m to 4 m dimensional block sizes but became significantly positively associated from 8 m sized blocks on. On the other hand the shrub species pairs; Lespedeza intermedia/ Robinia Pseudoacacia, and Lespedeza bicolor var, japonica/ Lespedeza intermedia were also significantly negatively associated from 1 m to 8 m sized blocks but became significantly positively associated from 16 m sized blocks on.
The associational pattern between Rhododendron mucronul tum var. poukhanenae and Lespedeza intermedia was not consistent throughout the stands. In some stands negative associations were observed throughout the blocks except NS 32.
From those observations micro-edaphic variation within the pine stands seems not to be great enough to cause distributional difference of pine trees within the ten pine stands. Among each shrub species and pine seedings, however, the edaphic variation within the pine stands may be great enough to cause distributional variation.
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